Glyptosaurinae

Glyptosaurinae Temporal range: Campanian–Oligocene PreꞒ Ꞓ O S D C P T J K Pg N
Fossil of the glyptosaurine Helodermoides tuberculatus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Family:	Anguidae
Subfamily:	†Glyptosaurinae Marsh, 1872
Subgroups
†Odaxosaurus †Paraplacosauriops †Peltosaurus †Proxestops †Xestops †Glyptosaurini †Glyptosaurus †Helodermoides †Paraglyptosaurus †Placosaurus †Proglyptosaurus †Stenoplacosaurus †Melanosaurini †Arpadosaurus †Melanosaurus

Glyptosaurinae is an extinct subfamily of anguid lizards that lived in the Northern Hemisphere during the Late Cretaceous and the Paleogene.

Description

Glyptosaurines are known primarily from their osteoderms, scale-like pieces of bone that are embedded in the skin and cover much of their bodies. The shape and extent of the osteoderms in glyptosaurines are similar to those seen in an unrelated group of lizards called Monstersauria, which includes the living Gila monster and beaded lizard.^[1] The osteoderms of glyptosaurines are unusually complex, consisting of four distinct layers of bony tissue. These tissues may have derived from both the dermis (the lower layer of the skin) and the epidermis (the outer layer of the skin) during their development in the embryo. The tissue forming the outermost layer of glyptosaurine osteoderms is similar to tooth enamel and has even been given its own name, osteodermine.^[2]

Glyptosaurines have been split into the subgroups Melanosaurini and Glyptosaurini, although recent phylogenetic analyses show that Melanosaurini in its traditional sense is paraphyletic, representing an evolutionary grade of glytosaurines more basal ("primitive") than Glyptosaurini. Below is a cladogram from Conrad and Norell (2008) showing the interrelationships of glyptosaurines and their relationship to other anguid lizards:^[1]

Evolutionary history

The earliest known Glyptosaurines like Odaxosaurus are known from the Late Cretaceous (Campanian) of North America. They arrived in Europe at the beginning of the Eocene (Ypresian). Only a single species, Stenoplacosaurus mongoliensis dating to the Mid-Eocene, is known from Asia. Glyptosaurines declined after the Mid-Eocene, become extinct in Europe by the end of the period as part of the "Grande Coupure".^[3] The youngest known remains of glyptosaurines is Peltosaurus granulosus from the Monroe Creek and upper Sharps formations of Sharps Corner, South Dakota, dating to the mid Oligocene, between 27.4 and 26.4 million years ago.^[4] The extinction of glyptosaurines was likely due to a cooling climate during the Oligocene.^[5]

References

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Extinct squamate genera

• Kingdom: Animalia
• Phylum: Chordata
• Class: Reptilia
• Superorder: Lepidosauria

Iguania
Acrodonta Anqingosaurus Barbaturex Bharatagama Gueragama Xianglong Agamidae Indiagama Suratagama Pleurodonta / Iguanoidea Anolbanolis Suzanniwana Uquiasaurus Xihaina Corytophanidae Aciprion? Babibasiliscus Geiseltaliellus †Gobiguania Anchaurosaurus Ctenomastax Igua Isodontosaurus Polrussia Saichangurvel Temujinia Zapsosaurus Iguanidae Afairiguana Polychrotidae Armandisaurus Lapitiguana Pariguana Pristiguana Pumilia Phrynosomatidae Desertiguana Other extinct iguanians Brasiliguana Hoyalacerta? Huehuecuetzpalli? Swainiguanoides †Arretosauridae Arretosaurus Dornosaurus Ergiliinsaurus Khaichinguana †Priscagamidae Heterodontagama Mimeosaurus Phrynosomimus Pleurodontagama? Priscagama

†Polyglyphanodontia / Borioteiioidea

Chamopsiidae Chamops Gerontoseps Glyptogenys Harmodontosaurus Haptosphenus Leptochamops Meniscognathus Pelsochamops Socognathus Stypodontosaurus Trippenaculus Gilmoreteiidae / "macrocephalosaurines" Gilmoreteius Kuwajimalla Yechilacerta "Polyglyphanodontines" Bicuspidon Cherminsaurus Darchansaurus Distortodon Erdenetesaurus Polyglyphanodon Sineoamphisbaena? Other polyglyphanodontians Adamisaurus Funiusaurus Gobinatus Obamadon Olindalacerta? Paleochelco? Peneteius Tchingisaurus Tianyusaurus Tijubina? Tripennaculus

Stem Scincogekkonomorpha
Scincogekkonomorpha Ardeosaurus? Bavarisaurus? Eichstaettisaurus? Jucaraseps Liushusaurus? Pedrerasaurus Scandensia? Yabeinosaurus? Scleroglossa Gekkonomorpha Norellius Gekkota Cadurcogekko Cretaceogekko Gobekko Hoburogekko Yanatarogecko Stem Autarchoglossa (Evansauria) Parmeosaurus Tepexisaurus Tijubina? †Bainguidae? Bainguis? Eoxanta Globaura Myrmecodaptria

Scincomorpha

Cordyloidea Eocordyla Ornatocephalus Sakurasaurus Lacertoidea Cnodontosaurus Paracontogenys Purbicella †Polyglyphanodontia? Amphisbaenia Oligodontosaurus Todrasaurus Amphisbaenidae Listromycter Lophocranion Bipedidae Anniealexandria Blanidae Blanosaurus Cuvieribaena Louisamphisbaena †Chthonophidae Chthonophis Rhineuridae Archaerhineura Dyticonastis Hadrorhineura Hyporhina Jepsibaena Macrorhineura Ototriton Plesiorhineura Protorhineura Spathorhynchus †Polyodontobaenidae Polyodontobaena †Contogeniidae Contogenys Palaeoscincosaurus Utahgenys Lacertidae Amblyolacerta Camptognathosaurus? Dormaalisaurus Dracaenosaurus Escampcerta Gracilicerta Ligerosaurus Maioricalacerta Mediolacerta Plesiolacerta Pseudeumeces Pseudolacerta Quercycerta Teiidae Brevisaurus? Meyasaurus? Ptilotodon †Barbatteiidae Barbatteius Oardasaurus Xantusiidae Catactegenys Palepidophyma Palaeoxantusia Scincoidea Retinosaurus Scincoideus Slavoia Scincidae Axonoscincus Orthoscincus Pyrenasaurus? Other scincomorphs Apsgnathus Calanguban Chromatogenys Dakotaseps Kleskunsaurus Lonchisaurus Monocnemodon Pachygenys Saurillodon Saurillus Sciroseps Webbsaurus †Paramacellodidae Atokasaurus? Becklesius Mimobecklesisaurus Paramacellodus Parasaurillus? Pseudosaurillus? Sharovisaurus

Anguimorpha
Basal anguimorphs Chometokadmon Eolacerta Paramacellodus Paravaranus? Yabeinosaurus? †Mosasauroidea? †Aigialosauridae †Mosasauridae Neoanguimorpha Carusioidea Carusia Xenosauridae Entomophontes Exostinus Restes Anguidae Odaxosaurus Anguinae Headonhillia Helvetisaurus Ophisauriscus Ragesaurus Diploglossinae Apodosauriscus Eodiploglossus Gerrhonotinae Paragerrhonotus Parophisaurus †Glyptosaurinae Arpadosaurus Melanosaurus Gaultia Glyptosaurus Helodermoides Odaxosaurus Paraglyptosaurus Paraplacosauriops Peltosaurus Placosaurus Proglyptosaurus Proxestops Xestops Paleoanguimorpha Basal varanoids / platynotans Aiolosaurus Colpodontosaurus Dorsetisaurus Eosaniwa Palaeovaranus Parasaniwa Parviderma Proplatynotia Saniwides Telmasaurus Shinisauria? Bahndwivici Dalinghosaurus Merkurosaurus Provaranosaurus Goannasauria Basal Goannasaurians Asprosaurus Chianghsia Estesia Eurheloderma Gobiderma Lowesaurus Morohasaurus Palaeosaniwa Paraderma Primaderma Varanoidea Lanthanotidae Cherminotus Varanidae Ovoo Saniwa Archaeovaranus Other platynotans / varanoids Arcanosaurus? Cemeterius Dryadissector

Ophidia
Coniophis Diablophis Dinilysia? Eophis Lapparentophis Najash? Parviraptor Portugalophis Tetrapodophis See Serpentes (snakes)

Indeterminate squamates
Costasaurus Lamiasaura Oculudentavis? Schillerosaurus

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